"You couldn't get to people for a long time on the island because roads were washed out, power was gone and the casualties mounted for a long time", Ryan said when asked about Trump's tweet.
She added another tweet saying the U.S. leader was "delusional, paranoid and unhinged from any sense of reality".
While Florence continued to ravage the Carolinas on Sunday, Long was forced to spend much of his time on "Meet the Press" answering questions about the investigation.
Researchers from the Milken Institute School of Public Health at George Washington University said the original estimates were so low because doctors on the island had not been trained to properly classify deaths after a natural disaster.
Trump called the federal response to Hurricane Maria an "unsung success" earlier in the week, and tweeted - without evidence - that the GWU study was an effort by Democrats to hurt him politically. Trump is so vain he thinks this is about him.
Roberto Firmino breaks silence after horrific eye injury v Tottenham
He was not even at his best for Watford against Manchester United, but was still one of the most impressive players on the park. We previously achieved it in both 1978-79 and 1990-91.
Boston Red Sox manager Alex Cora wasn't a big fan of President Donald Trump's comments Thursday about Puerto Rico. So when did people start dying?
A clear pattern emerged from the analysis: The mortality rate spiked in the months after the storm, particularly in the poorest areas of Puerto Rico, and among elderly males. "The president has done an extraordinary job in his operation FEMA and elsewhere going in there and rebuilding an island. rebuilding an electrical structure and everything else".
Other than from politicians, scorn flowed in from celebrities, the media, and the anti-Trump #resistance.
"We need to deal with the facts, which is these 2,975 people did not die of old age. We try to reach the truth, and we try to understand what is damaging the people in order to prevent disease", he said. "When you don't have food, you don't have water, no communication, no medicine".
San Juan Mayor Carmen Yulin Cruz, who has been in a war of words with Trump over the federal response to Hurricane Maria, shot back minutes later Thursday morning.
"This is what denial following neglect looks like", said Ms Cruz.
"The world has seen and the majority of the American people have seen how neglectful he was towards the people of Puerto Rico". Your lack of respect is appalling.
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