Clinton campaign manager John Podesta's emails were also hacked, which contained details of infighting and aides' disparaging remarks about Mrs Clinton.
The Kremlin, meanwhile, denied anew that it tried to sway the election.
The dates and content of the messages described in the indictment between the US person in regular contact with senior members of Trump's presidential campaign and Guccifer 2.0 match verbatim two screenshots Stone published on his website in March of 2017 of the direct message exchange he had with Guccifer 2.0 on Twitter in 2016. There is no allegation in this indictment that any American citizen committed a crime.
By June 2016, the defendants began planning the release of tens of thousands of stolen emails and documents, the indictment alleges.
In addition to contact with WikiLeaks, several people reached out to Guccifer 2.0 in August 2016 soliciting access to the stolen documents, the indictment says.
The charges come as Mr Mueller continues to investigate potential coordination between Russian Federation and the Trump campaign to influence the presidential election.
The indictment also does not allege that any vote tallies were altered by hacking.
Third, Friday's move also appears to show that the Mueller team and top Department of Justice officials such as Deputy Attorney General Rod Rosenstein are unintimidated by political threats against them.
The White House echoed some of those sentiments.
George Clooney reported hurt in motorcycle crash in Italy
The 57-year-old was travelling to the set of his new TV show Catch 22 in Costa Corallina when a auto turned and struck him. Clooney was reportedly on his way to the set of a TV series he's directing, based on the Joseph Heller classic "Catch-22".
The Department of Justice said the defendants used the cryptocurrency Bitcoin as part of a "conspiracy to launder money in which the defendants laundered the equivalent of more than $95,000 by transferring the money that they used to purchase servers and to fund other costs related to their hacking activities".
In February, Mueller charged 13 Russians and three Russian companies in an elaborate conspiracy to interfere in the election. The 20 are four former Trump campaign and White House aides, three of whom have pleaded guilty to different crimes and agreed to co-operate, and 13 Russians accused of participating in a hidden but powerful social media campaign to sway US public opinion in the 2016 election. "There is no debate that the Russians meddled in the election".
The GRU, which answers to the Russian military's General Staff, is part of the state machine and its involvement would indicate that the orders to interfere in the U.S. election came from the very top.
"I love the United States", Trump continued.
A few hours before the indictments were announced, Trump described the Mueller investigation as a "rigged witch hunt" that is hurting the U.S. relationship with Russian Federation.
The indictment was announced at nearly exactly the moment that Trump rolled into the quadrangle of Windsor Castle to meet the awaiting Queen Elizabeth II in the symbolic highpoint of his visit to Britain. They plan to hold a one-on-one meeting in Helsinki.
At the same press conference, Trump appeared to directly call upon Russian intelligence services to hack the former secretary of state's email account and to publish any information they stole as a part of their cyber-espionage operations within the 2016 presidential elections.
"When we confront foreign interference in American elections, it is important for us to avoid thinking politically as Republicans or Democrats and instead to think patriotically as Americans", he said.
President Donald Trump (R) and Russia's President Vladimir Putin talk as they make their way to take the "family photo" during the Asia-Pacific Economic Cooperation (APEC) leaders' summit in the central Vietnamese city of Danang on November 11, 2017. "Anything you do, it's always going to be, 'Oh, Russia, he loves Russia"'. "I did it. You got me.' There won't be a Perry Mason here", he said.
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